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During the maturation stage, the biofilm forms a complex 3D structure with nutrient and waste channels, which is sustained by continued EPS production and regulated by quorum sensing

Briefly, an aliquot of cell homogenate was incubated with the labeled substrate DEVD-pNA (acetyl-Asp-Glu-Val-Asp p-nitroaniline), which is cleaved by the caspase-3 enzyme and releases the chromophore pNA

The citric acid metabolites speak to mitochondrial health, with multiple markers inclusive of succinic and pyroglutamic acids pointing to glutathione status as a proxy for oxidative stress

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