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While we cannot rule out the existence of other substrates, Clade 2 GS-like enzymes have retained canonical activity

It downregulates Sterol Regulatory Element-Binding Protein (SREBP)-1c, Fatty Acid Synthase (FAS), Stearoyl-CoA Desaturase (SCD)1, Phosphoenolpyruvate Carboxykinase (PEPCK), Glucose-6-Phosphatase (G6Pase), and Apolipoprotein (Apo) C-III, while upregulating PPAR, Carnitine Palmitoyltransferase (CPT) 1, Acyl-CoA Dehydrogenase, Short Chain (ACADS), Pyruvate Dehydrogenase (PDase), Glycogen Synthase Kinase (GSK) 3, and Apo C-II (Ma et al., 2013

2016;167(4):1125-1136.e8

FoxO3 coordinates metabolic pathways to maintain redox balance in neural stem cells