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Acknowledgments This study was funded by the grants sponsored by the Ministry of Science and Technology: MOST 105-2313-B-033-001-MY3 and MOST 107-2622-B-033-001-CC2 to C.-D.H., and MOST108-2113-M153-003 to K.H.-C.C

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Free radicals are generated through several different pathways, e.g., during mitochondrial aerobic respiration, approximately 12% of electrons may escape the electron transport chain, and an estimated 2% of the oxygen consumed by mitochondria contributes to ROS formation, primarily the superoxide radical [25]