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However, under high-lactate conditions, lactate competes with the traditional substrate L-alanine for binding to the active site of AARS1, allowing it to operate as a lactyltransferase (23,95)

Concurrently, next-generation intelligent nanocarriers designed to synergistically induce ferroptosis and cuproptosis in response to tumor-specific stimuli should be developed to achieve spatiotemporal synergy