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Additionally, several studies have demonstrated the involvement of certain miRNAs in various GC activities [94], for instance, proliferation [66, 95,96,97], survival [98,99,100], terminal differentiation [101], steroidogenesis [96, 102,103,104], with the growth of the cumulus in mammals [105]

Besides, cytosol-mitochondria lactate shuttle also elaborates the role of L-lactate as the substrate source of mitochondrial ATP production (53, 5658)